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HCS612 Forage Crops
Adaption of Forage Species to Drought

Definitions

Drought

  • Meteorological
  • Biological
  • Sociological
Drought is a period of water deficit resulting in a negative impact on society (it is a human phenomenon)
Drought resistance - a generic term applied to tolerance and adaptation; with the exceptions of "hydraulic lift" and boundary layer effects, plants have little ability to resist drying by the atmosphere
Drought tolerance - the ability of plants to survive and grow during periods of water deficit
Drought adaptation - the ability of plants to modify their function to improve survival and growth during periods of water deficit
Drought avoidance - the ability of plants to improve survival and growth by escaping periods of water deficit (e.g. in a dormant state)
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Scales of Response

Community

  • sunny vs shady slopes
  • wet vs dry soils

tall grass prairie, eastern Nebraska
Big blue stem, switchgrass, indiangrass

short-grass prairie, western Nebraska
Blue grama, needle-and-thread

Morphological

  • Leaf loss
  • Smaller leaves (reduces the radiation load and transpiring area)
  • Reduced shoot growth
  • Greater absolute and/or relative root growth
  • Leaf senescence and death
  • transpiration (cools the leaf)

Anatomical and physiological

  • Concentration by dehydration
  • Osmotic adjustment
  • Proline accumulation
  • ABA increase
  • C3 vs C4 metabolism
  • Cell wall elasticity

Biochemical and molecular

  • Altered enzyme activity (a-amylase) is the result of molecular control
  • Heat shock proteins
  • Multiple genes
  • Water soluble carbohydrates (high sugar grasses)
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  Unique aspects of Forages

Pasture perenniality

  • Anticipated vs unexpected water deficit
  • Dilemma whether or not to continue growth into drought, water loss (ET) µ growth rate (photosynthesis)

Defoliation

  • Close grazing usually occurs during drought, but impairs recovery from drought
  • Effects of sward height and mass are unclear. Some studies show water conservation with long grass (mulch effect) – some show conservation with short grass (low transpiring surface) - probably a unimodal response confounded by dead matter
  • defoliation reduces root growth

Dry/severe environment

  • grasslands are usually in the more severe environments

Endophyte

  • unique to forage grasses

Quality

  • text 91-92
  • green:dead ratio
  • Loss of legumes
  • Increased fiber, reduced protein
  • Anti-quality

 

 

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